∙This study was financially supported by the Science Foundation of Hengyang Normal University (No. 19QD18), the Scientific Research Fund of Hunan Provincial Education Department (No. 19B083 and No. 20B084), The National Natural Science Foundation of China (No. 41671475), the Hunan Provincial Natural Science Foundation of China (No. 2019JJ50010), the Open Fund Project of Hunan Key Laboratory for Conservation and Utilization of Biological Resources in the Nanyue Mountainous Region (2021HSKFJJ033and NY20K03), and the Aid Program of Research Center for Environmental Education of Hunan Province (No. HJ19K02).