Conceptualization: L.J. and B.Z.; Data curation: X.C. and Y.J.; Formal analysis: L.J. and X.C.; Funding acquisition: B.Z.; Project administration: B.Z.; Validation: L.J. and X.C.; Visualization: Y.J.; Writing—original draft: L.J., X.C. and B.Z; Writing—review and editing: L.J. and B.Z. All authors have read and agreed to the published version of the manuscript. This research is funded by the National Natural Science Foundation of China (41901170, 41971183), the Natural Science Foundation of Guangdong Province (2018B030312004), Guangdong Basic and Applied Basic Research Foundation (2019A1515011385), and the Innovation Group Project of Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai) (311021018). All authors have read and agreed to the published version of the manuscript.