Conceptualization, L.Z. and R.C.; methodology, L.Z.; software, G.L.; validation, L.Z.; data curation, Y.Y.; writing—original draft preparation, L.Z.; writing—review and editing, Y.Y. and X.W.; funding acquisition, R.C. All authors have read and agreed to the published version of the manuscript. This research was funded by the Joint Research Project of Three-River Headwaters National Park, Chinese Academy of Sciences and the People’s Government of Qinghai Province (LHZX-2020-11), the National Natural Sciences Foundation of China (42171145), the Key Talent Program of Gansu Province, and the Gansu Provincial Science and Technology Program (22ZD6FA005).