Key Project of Hunan Provincial Education Department of China and Recruitment Program of High-end Foreign Experts of the State Administration of Foreign Experts Affairs of China funded this work via grant 17A067 and grant GWD201543000243. The China Scholarship Council Recruitment funded this work via grant 201808430277. The Talent Introduction Project of Hengyang Normal University also supported this work via grant 17D03. The United States National Science Foundation’s Division of Environmental Biology supported this work via grant OCE-1637630 for the Plum Island Ecosystems Long Term Ecological Research site. Opinions, findings, conclusions, and recommendations in this article are those of the authors and do not necessarily reflect those of the funders.